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主演：巴比·洛克伍德 巴瑞·波斯威克 萨缪尔·韦斯特 凯斯·索西 苏珊妮·布莱克史丽 杰夫·贝内特 莫里斯·拉马奇 杰森·亚历山大 马丁·肖特 蒂姆·本廷克 裘蒂·班森 玛丽·麦克劳德 本·蒂伯 伊利·罗素·林兹 卡莎·克罗宾斯基 迈克尔·勒纳 塔拉·斯特朗 凯瑟琳·博蒙特 吉姆·卡明斯
帕奇（巴比·洛克伍德 Bobby Lockwood 配音）和家人伙伴们来到了车水马龙的繁华大都市伦敦，这里的一切都令帕奇感到新奇无比，在不知不觉中，帕奇竟然和家人们走散了，它很快就被淹没在了川流不息的车流和人流之中。 在伦敦，孤身一人的帕奇遇见了名为雷电（巴瑞·波斯威克 Barry Bostwick 饰）的狗狗，这可乐坏了帕奇，因为雷电是经常在电视上出现，风靡全球的明星狗狗，两只狗很快就成为了好友，结伴而行。在经历了上一次的失败后，贪婪而又邪恶的克鲁拉（苏珊·布莱克利 Susan Blakely 饰）并没有放弃她的阴谋，这一次，她联手名为拉斯（马丁·肖特 Martin Short 饰）的艺术家，企图用斑点狗的皮来制作艺术作品。Roger, Anita and their canine clan are packing for the big move to Dalmatian Plantation, a home in the country with plenty of room for the 101 pups and far from the clutches of Cruella DeVil. The feistiest pup, Patch, feels lost in a sea of spots and longs to be a one-of-a-kind wonderdog like his TV hero, Thunderbolt. While watching the Thunderbolt Adventure Hour, Patch hears about a chance to appear on the show while it's filming in London. However, the family move will interfere with Patch's opportunity - until he's accidentally left behind in the commotion. Patch heads for the audition to meet his hero. Meanwhile, Thunderbolt's "trusty" sidekick, Lightning, reveals that the producers of the show want to replace Thunderbolt with a younger dog. In order to save his job, Thunderbolt decides he will go into the real world and perform an act of true heroism to prove himself. A veritable reference book to Thunderbolt's many adventures, Patch provides the perfect guide for the TV star in his attempts at real-life heroics. As for Cruella DeVil - she's back and more obsessed with Dalmatians than ever. At first, she's able to calm her frenzy through an affiliation with a spot-fixated artist.